Australoheros Rican & Kullander, 2006
type species: Australoheros facetus (Jenyns, 1842)
Australoheros tembe (Casciotta et al., 1995)
Australoheros scitulus (Rican & Kullander, 2003)
Finally the ‘Cichlasoma’-facetum-group was defined formally as a genus: Australoheros.
Since this group was taken out from the real Cichlasoma by Kullander in 1983 it was referred to as ‘Cichlasoma’-facetum-group containing: ‘Cichlasoma’ facetum, ‘C’. tembe and ‘C’. scitulum.
The new genus consists the three species mentioned above and at least further seven undescribed species: Australoheros sp. ‘Forquilha’, A. cf. facetus, A. sp. ‘Jacui’, A. sp. ‘Jacutinga’, A. sp. ‘Paraguay’, A. sp. ‘Pirapo’, and Australoheros sp. ‘Uruguai’.
These seven are temporary references sensu Art. 1.3.5 by avoiding binominal names, using ‘cf.’ in A. cf. facetus and combinations of genus, sp. and the location name of the specimens in the other six. The rules for available names are not to be applied.
Within the discussion part of the paper the authors probably by mistake mention A. sp. jacutinga as Australoheros jacutinga, a binominal name printed in italics. Thus Australoheros jacutinga might be a nomen nudum in the sense of the ICZN.
Rican, O. & S. Kullander (2006):
Character- and tree-based delimitation of species in the ‘Cichlasoma’ facetum group (Teleostei, Cichlidae) with the description of a new genus.
Journal of Zoological Systematics and Evolutionary Research 44 (2): 136-152
abstract (from publication):
The ‘Cichlasoma’ facetum group is part of the taxonomically complex group of Neotropical cichlid fishes of the tribe Heroini. Many species groups and unplaced species of heroines are still left without a generic name following the revision of the genus Cichlasoma.
We describe here the ‘Cichlasoma’ facetum group as a new genus, Australoheros, and provide evidence for its monophyly based on phylogenetic analyses of morphological and mtDNA characters. Australoheros is morphologically characterized by the lowest values in meristic characters among heroines and by three apomorphic characters in coloration pattern. In addition to the three described species of Australoheros, our results of species delimitation based on a combination of tree- and character-based approaches identify seven putatively new species of Australoheros.
Several coding schemes of morphological characters are used to recover the intrageneric relationships within the genus, resulting in very similar topologies. Discovery of additional species within the genus is expected once material from the whole distribution area is studied.