Loricaria holmbergi – first record from Paraguay

first record from Paraguay:
Loricaria holmbergi Rodriguez & Miquelarena, 2005

published in:
Londoño-Burbano, A., A. Urbano-Bonilla & M.R. Thomas (2020):
Loricaria cuffyi (Siluriformes: Loricariidae), a new species of loricariin catfish from the Guiana Shield.
Journal of Fish Biology (doi:10.1111/jfb.14566, preprint: 1-4)

abstract (from publication):
Loricaria cuffyi n. sp. is described based on 36 specimens from the Essequibo and upper Negro River drainages in western Guyana and the upper Orinoco River drainage in Venezuela. The new species can be distinguished from sympatric and geographically proximate congeners by a postorbital notch that is inconspicuous, shallow and rounded, odontode ridges on the dorsum of head and predorsal weakly developed, abdominal plates tightly joined and completely covering the median abdominal space and pectoral girdle, higher anterior lateral plate counts, and coloration characteristics.
The distribution of the new species adds to an interesting and welldocumented biogeographical pattern exhibited by other Guiana Shield loricariids influenced by the proto-Berbice during the Cenozoic and recent configuration of drainages in the Guiana Shield. We present an update on the taxonomy of Loricaria, and discuss the biogeography and conservation status of the new species.